Differentiability with respect to a parameter for a Lotka-Volterra system with delays, via step method

Abstract

In this paper, using the step method, we establish the differentiability with respect to parameter for a Lotka-Volterra system with two delays.

Authors

Diana Otrocol

Diana Otrocol

Tiberiu Popoviciu Institute of Numerical Analysis Romanian Academy

Keywords

Differential equations; delay, step method

Paper coordinates

D. Otrocol, Differentiability with respect to a parameter for a Lotka-Volterra system with delays, via step method, Rev. Anal. Numér. Théor. Approx., 35 (2006), no. 1, 83-86.

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Rev. Anal. Numér. Théor. Approx.

Publisher Name

Romanian Academy

Print ISSN

 2457-6794

Online ISSN

2501-059X

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[1] Hale, J., Functional differential equations, Springer-Verlag, New York, Heidelberg, Berlin, 1977.
[2] Hale, J. and Ladeira, L., Differentiability with respect to delays, J. Differential Equations, 92, pp. 14–26, 1991.
[3] Hartman, P., Ordinary differential equations, John Wiley & Sons, Inc., New York, London, Sydney, 1964.
[4] Hokkanen, V. M. and Morosanu, Gh., Differentiability with respect to delay, Differential Integral Equations, 11, no. 4, pp. 589–603, 1998.
[5] Mures¸an, V., Differential equations with linear modified argument, Ph. D. Thesis, “Babes-Bolyai” University, 1997.
[6] Otrocol, D., Data dependence for the solution of a Lotka-Volterra system with two delays, Mathematica (to appear).
[7] Otrocol, D., Smooth dependence on parameters for some Lotka-Volterra system with delays (to appear).
[8] Rus, I. A., Generalized contractions, Seminar of Fixed Point Theory, “Babes-Bolyai” University, pp. 1–130, 1983.
[9] Rus, I. A., Functional-differential equation of mixed type, via weakly Picard operators, Seminar of Fixed Point Theory, Cluj-Napoca, 3, pp. 335–346, 2002.
[10] Rus, I. A., Generalized Contractions and Applications, Cluj University Press, 2001.
[11] Serban, M. A., Fiber ϕ-contractions, Studia Univ. “Babes-Bolyai”, Mathematica, 44, no. 3, pp. 99–108, 1999.
[12] Tamasan, Al.  , Note on Differentiability with respect to delay, P.U.M.A., 9, nos. 1–2, pp. 215–220, 1998.

DIFFERENTIABILITY WITH RESPECT TO A PARAMETER FOR A LOTKA-VOLTERRA SYSTEM WITH DELAYS, VIA STEP METHOD∗

DIANA OTROCOL†
(Received by the editors: ??)
Abstract.

In this paper, using the step method, we establish the differentiability with respect to parameter for a Lotka-Volterra system with two delays.

Keywords:
Differential equations, delay, step method.
1991 Mathematics Subject Classification:
34L05, 47H10.
∗This work has been supported by MEdC-ANCS under grant ET 3233/17.10.2005
†“Tiberiu Popoviciu" Institute of Numerical Analysis, P.O.Box. 68-1, Cluj-Napoca, Romania

1. INTRODUCTION

The purpose of this paper is to study the λ\lambda-dependence of the solution of the Lotka-Volterra problem:

(1) xi′​(t)=fi​(t,x1​(t),x2​(t),x1​(t−τ1),x2​(t−τ2);λ),t∈[t0,b],λ∈J\displaystyle x_{i}^{\prime}(t)=f_{i}(t,x_{1}(t),x_{2}(t),x_{1}(t-\tau_{1}),x_{2}(t-\tau_{2});\lambda),\ \ \ \ \ \ t\in[t_{0},b],\ \lambda\in J
(4) x1​(t)=φ​(t),t∈[t0−τ1,t0],x2​(t)=ψ​(t),t∈[t0−τ2,t0].\displaystyle\begin{array}[c]{c}x_{1}(t)=\varphi(t),\ t\in[t_{0}-\tau_{1},t_{0}],\\ x_{2}(t)=\psi(t),\ t\in[t_{0}-\tau_{2},t_{0}].\end{array}

There have been many studies on this subject. Differentiability with initial data for the functional differential equations was first established by Hale in [2], but differentiability with respect to delays for delay differential equations was proved by Hale and Ladeira in [3] and by A. Tămăşan in [13]. The paper of Hokkanen and Moroşanu [5] gives a proof for delay differential equations case using the step method. The Picard operators’ technique proposed by I.A. Rus [9], [10], [11], was used by V. Mureşanu [6] to prove continuity with respect to λ\lambda, M. Şerban [12], D. Otrocol [8] using the theorem of fibre contraction.

In this paper we use the following theorem for the simple case of ordinary differential system.

Theorem 1.

[4] Consider the initial value problem

x′​(t)=g​(t,x​(t);λ)x​(t)=x0,t∈[a,b]\begin{array}[c]{l}x^{\prime}(t)=g(t,x(t);\lambda)\\ x(t)=x_{0}\end{array},\ t\in[a,b]

where g∈C1​([a,b]×ℝ×J,ℝm),‖∂g​(t,u,λ)∂u‖ℝm≤M1g\in C^{1}([a,b]\times\mathbb{R}\times J,\mathbb{R}^{m}),\ \big\|\tfrac{\partial g(t,u,\lambda)}{\partial u}\big\|_{\mathbb{R}^{m}}\leq M_{1}. Then the unique solution x∈C1​([a,b]×J)x\in C^{1}([a,b]\times J).

2. MAIN RESULT

We consider the following Lotka-Volterra system with parameter:

(5) xi′​(t)=fi​(t,x1​(t),x2​(t),x1​(t−τ1),x2​(t−τ2);λ),t∈[t0,b],λ∈J\displaystyle x_{i}^{\prime}(t)=f_{i}(t,x_{1}(t),x_{2}(t),x_{1}(t-\tau_{1}),x_{2}(t-\tau_{2});\lambda),\ \ \ \ \ \ t\in[t_{0},b],\ \lambda\in J
(8) x1​(t)=φ​(t),t∈[t0−τ1,t0],x2​(t)=ψ​(t),t∈[t0−τ2,t0].\displaystyle\begin{array}[c]{c}x_{1}(t)=\varphi(t),\ t\in[t_{0}-\tau_{1},t_{0}],\\ x_{2}(t)=\psi(t),\ t\in[t_{0}-\tau_{2},t_{0}].\end{array}

We suppose that:

  1. (H1)

    τ1≤τ2;t0<b;J⊂ℝ\;\tau_{1}\leq\tau_{2};\;t_{0}<b;\;J\subset\mathbb{R}, a compact interval ;

  2. (H2)

    fi∈C1​([t0,b]×ℝ4×J),i=1,2\;f_{i}\in C^{1}([t_{0},b]\times\mathbb{R}^{4}\times J),\;i=1,2;

  3. (H3)

    ∃M>0\;\exists M>0 such as ‖∂fi∂uj​(t,u1,u2,u3,u4;λ)‖ℝn≤M1,\big\|\frac{\partial f_{i}}{\partial u_{j}}(t,u_{1},u_{2},u_{3},u_{4};\lambda)\big\|_{\mathbb{R}^{n}}\leq M_{1},\;\newline for allt∈[t0,b],uj∈Rn,j=1,4¯,λ∈J,i=1,2\ t\in[t_{0},b],\;u_{j}\in R^{n},\;j=\overline{1,4},\;\lambda\in J,\ i=1,2;

  4. (H4)

    φ∈C​([t0−τ1,t0]),ψ∈C​([t0−τ2,t0])\;\varphi\in C([t_{0}-\tau_{1},t_{0}]),\;\psi\in C([t_{0}-\tau_{2},t_{0}]).

In the above conditions, from Theorem 1 in [7] we have that the problem (5)–(8) has a unique solution, (x1​(t;λ),x2​(t;λ))(x_{1}(t;\lambda),x_{2}(t;\lambda)).

We prove that

x1​(t;⋅)\displaystyle x_{1}(t;\cdot) ∈C1​(J)​, for all ​t∈[t0−τ1,b],\displaystyle\in C^{1}(J)\text{, for all }t\in[t_{0}-\tau_{1},b],
x2​(t;⋅)\displaystyle x_{2}(t;\cdot) ∈C1​(J)​, for all ​t∈[t0−τ2,b].\displaystyle\in C^{1}(J)\text{, for all }t\in[t_{0}-\tau_{2},b].

applying the Theorem 1 and using the step method.

We consider the system

(9) xi′​(t,λ)=fi​(t,x1​(t,λ),x2​(t,λ),x1​(t−τ1,λ),x2​(t−τ2,λ);λ)t∈[t0,b],λ∈J\begin{array}[c]{c}x_{i}^{\prime}(t,\lambda)=f_{i}(t,x_{1}(t,\lambda),x_{2}(t,\lambda),x_{1}(t-\tau_{1},\lambda),x_{2}(t-\tau_{2},\lambda);\lambda)\\ t\in[t_{0},b],\ \lambda\in J\end{array}

with initial conditions

(10) x1​(t;λ)=φ​(t;λ)x2​(t;λ)=ψ​(t;λ)\begin{array}[c]{c}x_{1}(t;\lambda)=\varphi(t;\lambda)\\ x_{2}(t;\lambda)=\psi(t;\lambda)\end{array}

where x1∈C​([t0−τ1,b]×J)∩C1​[t0,b],x2∈C​([t0−τ2,b]×J)∩C1​[t0,b]x_{1}\in C([t_{0}-\tau_{1},b]\times J)\cap C^{1}[t_{0},b],\;x_{2}\in C([t_{0}-\tau_{2},b]\times J)\cap C^{1}[t_{0},b].

Theorem 2.

In conditions (H1), (H2), (H4), the solution (x1​(t;λ),x2​(t;λ))(x_{1}(t;\lambda),x_{2}(t;\lambda)) of the problem (9)–(10) is smooth continuosly with respect to λ\lambda.

Theorem 3.

In conditions(H1)–(H4), the solution (x1​(t;λ),x2​(t;λ))(x_{1}(t;\lambda),x_{2}(t;\lambda)) of the problem (9)–(10) is differentiable with respect to λ\lambda.

We prove that

x1​(t;⋅)\displaystyle x_{1}(t;\cdot) ∈C1​(J)​, for all ​t∈[t0−τ1,b],\displaystyle\in C^{1}(J)\text{, for all }t\in[t_{0}-\tau_{1},b],
x2​(t;⋅)\displaystyle x_{2}(t;\cdot) ∈C1​(J)​, for all ​t∈[t0−τ2,b].\displaystyle\in C^{1}(J)\text{, for all }t\in[t_{0}-\tau_{2},b].

applying the Theorem 1 and using the step method.

For t∈[t0,t0+τ1]t\in[t_{0},t_{0}+\tau_{1}], from the Theorem 1 we have that x1​(t;⋅)∈C1​(J),x_{1}(t;\cdot)\in C^{1}(J), x2​(t;⋅)∈C1​(J)x_{2}(t;\cdot)\in C^{1}(J).

For t∈[t0+τ1,t0+2​τ1]t\in[t_{0}+\tau_{1},t_{0}+2\tau_{1}], from the Theorem 1 we have that x1​(t;⋅)∈C1​(J),x2​(t;⋅)∈C1​(J)x_{1}(t;\cdot)\in C^{1}(J),\ x_{2}(t;\cdot)\in C^{1}(J). From (9) and (H2)(H_{2}) it follows that x1′​(t0+τ1−0)=x1′​(t0+τ1+0)x_{1}^{\prime}(t_{0}+\tau_{1}-0)=x_{1}^{\prime}(t_{0}+\tau_{1}+0) and x2′​(t0+τ1−0)=x2′​(t0+τ1+0)x_{2}^{\prime}(t_{0}+\tau_{1}-0)=x_{2}^{\prime}(t_{0}+\tau_{1}+0). Then (x1​(t;λ),x2​(t;λ))(x_{1}(t;\lambda),x_{2}(t;\lambda)) is differentiable in t0+τ1t_{0}+\tau_{1}.

For t∈[t0+n​τ1,t0+τ2]t\in[t_{0}+n\tau_{1},t_{0}+\tau_{2}], from the Theorem 1 we have that x1​(t;⋅)∈C1​(J),x2​(t;⋅)∈C1​(J)x_{1}(t;\cdot)\in C^{1}(J),\ x_{2}(t;\cdot)\in C^{1}(J). From (9) and (H2)(H_{2}) it follows that x1′​(t0+n​τ1−0)=x1′​(t0+n​τ1+0)x_{1}^{\prime}(t_{0}+n\tau_{1}-0)=x_{1}^{\prime}(t_{0}+n\tau_{1}+0) and x2′​(t0+n​τ1−0)=x2′​(t0+n​τ1+0)x_{2}^{\prime}(t_{0}+n\tau_{1}-0)=x_{2}^{\prime}(t_{0}+n\tau_{1}+0). Then (x1​(t;λ),x2​(t;λ))(x_{1}(t;\lambda),x_{2}(t;\lambda)) is differentiable in t0+n​τ1t_{0}+n\tau_{1}.

So (x1​(t;λ),x2​(t;λ))(x_{1}(t;\lambda),x_{2}(t;\lambda)) is C1C^{1} in the knots. ∎

We present below a simple example to illustrate the procedures of applying our results.

Example 1.

Consider the Lotka-Volterra type predator-prey system with two delays with parameter:

(11) x1′​(t,λ)=λ​x1​(t;λ)​[2−x1​(t;λ)−x2​(t;λ)]x2′​(t,λ)=λ​x2​(t;λ)​[2−x1​(t−1;λ)−x2​(t−2;λ)],t∈[0,2]\begin{array}[c]{l}x_{1}^{\prime}(t,\lambda)=\lambda x_{1}(t;\lambda)\left[2-x_{1}(t;\lambda)-x_{2}(t;\lambda)\right]\\ x_{2}^{\prime}(t,\lambda)=\lambda x_{2}(t;\lambda)\left[2-x_{1}(t-1;\lambda)-x_{2}(t-2;\lambda)\right]\end{array},\ t\in[0,2]

with initial conditions

(12) x1​(t;λ)=t+1,t∈[t0−1,t0],x2​(t;λ)=t+2,t∈[t0−2,t0].​\equation@qed\begin{array}[c]{c}x_{1}(t;\lambda)=t+1,\ t\in[t_{0}-1,t_{0}],\\ x_{2}(t;\lambda)=t+2,\ t\in[t_{0}-2,t_{0}].\end{array}\equation@qed

Equation (11) is the form of (9). Therefore, the assumptions (H1)−(H4)(H_{1})-(H_{4}) are satisfied for (11)–(12). Moreover, the Theorem 2 hold. Following the proof of Theorem 3 we see that:

For t∈[t0,t0+1]t\in[t_{0},t_{0}+1], from the Theorem 1 we have that x1​(t;⋅)∈C1​(J),x2​(t;⋅)∈C1​(J)x_{1}(t;\cdot)\in C^{1}(J),\ x_{2}(t;\cdot)\in C^{1}(J).

Refer to caption
Fig. 1.. Comparison between Lotka-Volterra systems with different parameters.

For t∈[t0+1,t0+2]t\in[t_{0}+1,t_{0}+2], from the Theorem 1 we have that x1​(t;⋅)∈C1​(J),x2​(t;⋅)∈C1​(J)x_{1}(t;\cdot)\in C^{1}(J),\ x_{2}(t;\cdot)\in C^{1}(J). From (9) and (H2)(H_{2}) it follows that x1′​(t0+1−0)=x1′​(t0+1+0)x_{1}^{\prime}(t_{0}+1-0)=x_{1}^{\prime}(t_{0}+1+0) and x2′​(t0+1−0)=x2′​(t0+1+0)x_{2}^{\prime}(t_{0}+1-0)=x_{2}^{\prime}(t_{0}+1+0). Then (x1​(t;λ),x2​(t;λ))(x_{1}(t;\lambda),x_{2}(t;\lambda)) is differentiable in t0+1t_{0}+1. So (x1​(t;λ),x2​(t;λ))(x_{1}(t;\lambda),x_{2}(t;\lambda)) is C1C^{1} in the knots. Therefore the Theorem 3 is proved.

Here we give a portrait of the trajectory of (11)–(12) drawn by a computer using MATLAB facilities. The results from numerical computation are plotted for λ=1.35,1.37,1.39,1.41\lambda=1.35,1.37,1.39,1.41 in Fig. 1.

References

  • [1]
  • [2] Hale, J., Functional differential equations, Springer Verlag, New-York, Heidelberg, Berlin, 1977.
  • [3] Hale, J. and Ladeira, L., Differentiability with respect to delays, J. Differential Equations, 92, pp. 14–26, 1991.
  • [4] Hartman, P., Ordinary differential equations, John Wiley & Sons, Inc., New-York, London, Sydney, 1964.
  • [5] Hokkanen, V. M. and Moroşanu, Gh., Differentiability with respect to delay, Differential Integral Equations, 11, no. 4, pp. 589–603, 1998.
  • [6] Mureşan, V. , Differential equations with linear modified argument, Ph. D. Thesis, “Babeş-Bolyai” University, 1997.
  • [7] Otrocol, D. , Data dependence for the solution of a Lotka-Volterra system with two delays, Mathematica, (to appear).
  • [8] Otrocol, D., Smooth dependence on parameters for some Lotka-Volterra system with delays, (to appear).
  • [9] Rus, I. A., Generalized contractions, Seminar of Fixed Point Theory, “Babeş-Bolyai” University, pp. 1–130, 1983.
  • [10] Rus, I. A., Functional-differential equation of mixed type, via weakly Picard operators, Seminar of Fixed Point Theory, Cluj-Napoca, 3, pp. 335–346, 2002.
  • [11] Rus, I. A., Generalized Contractions and Applications, Cluj University Press, 2001.
  • [12] Şerban, M. A., Fiber φ\varphi-contractions, Studia Univ. “Babeş-Bolyai”, Mathematica, 44, no. 3, pp. 99–108, 1999.
  • [13] Tămăşan, Al., Note on Differentiability with respect to delay, P.U.M.A., 9, nos. 1–2, pp. 215–220, 1998.
  • [14]
2006

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